GESSEL, S. P. and TURNER, J.: Litter production by red alder in western Washington. For. Sci. 20: 325-330, 1974
KIKUZAWA, K.: Emergence, defoliation and longevity of alder (Alnus hirsuta TURCZ.) leaves in a deciduous hardwood forest stand. Jpn. J. Ecol. 28: 299-306, 1978
菊沢喜八郎:ハンノキ属の葉はなぜ夏に落ちるか.日生態会誌 30: 359~368, 1980
KIKUZAWA, K., ASAI, T. and FUKUCHI, M.: Leaf-litter production in a plantation of Alnus inokumae. Jpn. J. Ecol. 72: 993-999, 1984