Research field (3):
Evolutionary biology
, Animals: biochemistry, physiology, behavioral science
, Ecology and environmental science
European Society for Evolutionary Biology
, 日本生態学会
, International Society for Behavioral Ecology
, 日本進化学会
, 日本鳥学会
, 日本動物行動学会
, European Society for Evolutionary Biology
, Japan Ecological Society
, Society for Evolutionary Studies, Japan
, The Ornithological Society of Japan
, Japan Ethological Society
, International Society for Behavioral Ecology