日本近化学会
, SOCIETY FOR THE STUDY OF AMRHIBIANS AND REPTILES
, AMERICAN SOCIETY OF ICHTHYOLOGISTS AND HERPETOLOGISTS
, 動物分類学会
, 日本爬虫両棲類学会
, 日本動物学会
, Society fro the Study of Amphibians and Reptiles
, American Society of Ichthyologists and Herpetologists
, Society of Evolutionary Studies, Japan
, Biogeographical Society of Japan
, Japanese Society of Systematic Zoology
, Zoological Society of Japan
, Herptological Society of Japan