BUCHLI, H. L'origine des castes et les potentialites ontogeniques des termites europeens du genre Reticulitermes Holmgren. Annales Des Sciences Naturelles, Zoologie, 11e Serie. 1958, 20, 263-429
CHIUH, T. Notes on the types of the reproductive castes of subterranean termites, Coptotermes formosanus Shiraki and Reticulitermes flaviceps Oshima (Isoptera : Rhinotermitidae). Acta Entomologica Sinica. 1960, 10, 302-306
HAMILTON, WD. Extraordinary sex ratios. Science. 1967, 156, 477-488
HARE, L. Caste determination and differentiation with special reference to the genus Reticulitermes (Isoptera). Journal of Morphology. 1934, 56, 267-293
HOWARD, RW. Reproductives in mature colonies of Reticulitermes flavipes : abundance, sex-ratio and association with soldiers. Environmental Entomology. 1980, 9, 458-460