Abe, A, Kudoh, K. & Saitoh, K., 1976. Chromosome study in genera Prosopocoilus, Nipponodorcus and Macrodorcas of Lucanidae. Science Reports of Hirosaki University, 23: 50-56.
Abe, A. & Kudoh, K., 2005. Chromosomes of three species of Dorcus (Coleoptera, Lucanidae), with a note on their sex chromosome evolution. Elytra, Tokyo, 33: 513-521.
阿部東・工藤貢次・寺山博, 2006. クワガタムシ科 Dorcus 7種の染色体調査. kogane, Tokyo, (7): 87-98.