Kiyotaka Hori, Daiki Takahashi, Katsuhiro Yoneoka, Ying-Hong He, Yoshihisa Suyama, Noriaki Murakami, Atsushi Ebihara. Arachniodes hiugana (Dryopteridaceae) is a Diploid Hybrid. Bulletin of the National Museum of Nature and Science Series B (Botany). 2025. 51. 4. 137-146
堀清鷹. Origin of Japanese Woodsia intermedia (Woodsiaceae, Polypodiiae) based on phylogenetic analyses of plastid rbcL and nuclear pgiC genes. Hikobia. 2024. 19. 61-65