- 1997 - 1999 Heidelberg University
- 1997 - 1999 Biochemie Zentrum,University of Heidelberg
- 1999 - - 現職
- 1999 - - Research Institute for microbial Diseases,Osaka
- 1994 - 1997 Osaka University Research Institute for Microbial Diseases
- 1994 - 1997 Research Institute for Microbial Diseases,Osaka
- 1990 - 1994 大阪府立大学農学研究科博士後期課程
- 1990 - 1994 Faculty of Agriculture,Osaka Prefecture University
- 1989 - 1990 日本チバガイギー(株)薬理学研究室研究員
- 1989 - 1990 Research Scientist at the Dept,Pharmacology
- 1987 - 1989 大阪府立大学農学研究科博士前期課程
- 1987 - 1989 Faculty of Agriculture,Osaka Prefecture University
- Kyoto University Institute for Virus Research
- UNiversity
- Institute for Virus Research,Kyoto University
- CIBA-GEIGY,Japan
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日本細胞生物学会
, 日本RNA学会
, 日本分子生物学会
, Molecular Biology Society Japan
, Japanese Society for Microbiology