1979 - 1983 Osaka University Graduate School, Division of Medicine
1973 - 1979 The University of Tokyo Faculty of Medicine
Professional career (1)：
M. D. and Ph. D. (Osaka University)
Work history (5)：
2000/09/01 - 2016/09/30 Hiroshima University Professor
1990/07/01 - 2000/08/31 Tokyo Metropolitan Institute for Neuroscience Head Researcher
1985/04/01 - 1990/06/30 Kyoto University Assistant Professor
1987/06/16 - 1990/06/08 Carnegie Institution of Washington Research Assistant
1983/10/01 - 1985/03/31 Osaka University Assistant Professor
Association Membership(s) (4)：
The Japanese Society of Developmental Biologists
, The Moleclar Biology Society of Japan
, The Japanese Biochemical Socity
, the Zoological Society of Japan
