Research field (3):
Biodiversity and systematics
, Animals: biochemistry, physiology, behavioral science
, Ecology and environmental science
日本野生生物保護学会
, 日本生態学会
, 日本昆虫学会
, 日本土壌動物研究会
, 日本蟻類研究会
, 国際社会性昆虫学会(International union for the study of social Insects)
, The ecological society of Japan
, The entomological society of Japan
, The Japanese society of soil zoology
, The myrmecological society of Japan
, International union for the study of social insects