- 2018 - 2021 Sperm recognition mechanisms and matrix formation mechanisms of mammalian egg coats
- 2015 - 2018 Sperm recognition domain in mammalian egg coat matrix
- 2012 - 2015 Sperm recognition domain in mammalian egg coat matrix
- 2009 - 2011 Sperm recognition and matrix formation mechanisms of mammalian egg coats
- 2007 - 2008 Structure and function of mammalian egg zona pellucida : are the sugarchains really essential for zona functions
- 2006 - 2007 Structural and Functional Analysis of glycoproteins participating in the mammalian fertilization and development
- 2001 - 2002 哺乳動物の生殖反応マシーナリーの構造機能解析
- 1999 - 2000 哺乳類受精における多精拒否へのシアル酸およびシアリダーゼの関与
- 1998 - 2000 Structure and Function of Cell Surface Molecules
- 1994 - 1995 Structure and Function of Pig and Bovine Egg Zona Pellucida
- 1994 - 1994 哺乳類卵子透明帯の精子結合部位に関する解析
- 1993 - 1994 ブタ精子の卵子透明帯との結合の分子機構
- 1992 - 1992 ブタ卵子透明帯糖タンパク質の精子結合部位の解析
- 1990 - 1992 Functional analysis of mammalian actin regulatory proteins, cofilin and destrin
- 1991 - 1991 哺乳類の受精における多精拒否機構の研究
- 1991 - 1991 ブタ卵子透明帯糖タンパク質の遺伝子の単離と構造解析
- 1989 - 1990 アクチン結合蛋白質コフィリンの構造と機能
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