- 2021 - 2024 半クローン雑種の総括研究-永続性の実証(究極要因)とゲノム削除機構(至近要因)
- 2017 - 2020 Origin and evolution of hemiclonal Hexagrammos hybrids elongating the lineage lifetime by host-switching
- 2016 - 2019 Evolutionary and ecological studies on sperm and reproductive modes in marine sculpins
- 2015 - 2019 Molecular mechanisms of atypical development and reproduction caused by hybridization and the application to breeding in fishes
- 2014 - 2017 Change of cold currents fish fauna by expansion of warm currents - migration, settlement or genome-parasitism
- 2013 - 2016 Parapatric evolution and adaptive divergence of Insular fishes along the Oyashio current
- 2011 - 2014 Genetic and ecological studies of the origin and reproduction about hemi-clonal hybrids that were first found from marine fishes
- 2010 - 2012 Study on the reproductive behavior and the sociality in Lake Tanganyika cichlids as a model system of cooperative breeding in vertebrates
- 2009 - 2012 Study on terrific adaptive radiation of Baikal sculpins by biodiversity of the breeding ecology, early life history and genes
- 2007 - 2008 Quantitative estimation of sperm competition in the evolution of thereproductive styles in cottoid fishes as a model of copulatory animals
- 2005 - 2007 A study of reproductive ecology of Lake Tanganyikan cichlid fishes as the model systems of cooperative breeding in vertebrates
- 2005 - 2007 Evolutionary and ecological studies on the circum-north Pacific elements, the super-family cottoid
- 2004 - 2006 Ecological Studies on Crossbreeding among Hexagrammos Species
- 2004 - 2006 A study on reproductive tactics in four species of cichlid fishes: model systems of cooperative breeding in vertebrates
- 2004 - 2005 スキューバ潜水による着底稚魚の効率的採集機材の開発
- 2002 - 2004 Study of reproductive ecology and sociality of Lake Tanganyika cichlids : a model system of social evolution of vertebrates
- 2001 - 2003 Study on social and ecological factors promoting cooperative polyandry in fish and on their reproductive tactics
- 2001 - 2002 Studies on the diversity and creative mechanism of the reproductive mode in copulating teleost fishes
- 1998 - 2000 Studies on the reproductive ecology of littoral animals using with DNA polymorphism
- 1998 - 1998 魚類における温度依存性決定機構の生態的意義
- 1996 - 1997 Survey for resource estimation of substrate spawning littoral fishes with using underwater video camera system
- 1995 - 1995 沿岸性魚類の卵保護習性に関する資源生態学的研究
- 1993 - 1993 カジカ科魚類における交尾の成立基盤
- 1989 - 1990 カサゴ目魚類における生殖様式の進化 交尾型カジカの行動様式と受精について
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