International Association for Plant Taxonomy
, International Association of Bryologists
, The Botanical Society of Chugoku-Shikoku
, HIKOBIA BOTANICAL SOCIETY
, THE BRYOLOGICAL SOCIETY OF JAPAN
, SOCIETY OF EVOLUTIONARY STUDIES, JAPAN
, THE JAPANESE SOCIETY OF PLANT PHYSIOLOGISTS
, JAPAN SOCIETY FOR CELL BIOLOGY
, JAPANESE SOCIETY FOR PLANT SYSTEMATICS
, THE JAPANESE SOCIETY OF PLANT MORPHOLOGY
, THE BOTANICAL SOCIETY OF JAPAN