- 2024 - 2029 チビゴキブリ科の発生学的研究-ゴキブリ目、網翅類の系統学的再構築を目指して-
- 2020 - 2025 Evolutionary scenario of adaptation to terrestrial environment in Insect
- 2019 - 2024 Diversity of fertilization in the basal lineages of insects, for deepening our insight to insect early evolution
- 2016 - 2020 Embryological study on proturan brain: groudplan and evolution of Hexapoda
- 2016 - 2019 Diversity of fertilization in the basal lineages of insects, for deepening our insight to insect early evolution
- 2007 - 2010 サイコクカマアシムシの発生学的研究-六脚類の高次系統・グラウンドプランの再構築-
- 2007 - 2009 サイコクカマアシムシの発生学的研究-六脚類の高次系統・グラウンドプランの再構築-
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