- 2022/04 - 現在 日本学術振興会 RPD 特別研究員
- 2018/04 - 現在 JSPS RPD researcher
- 2017/04 - 現在 Kyoto University Graduate school of Biostudies Post doctoral researcher
- 2010/07 - 2017/03 Kyoto University iCeMS Post doctoral researcher
- 2012/04 - 2015/03 JSPS PD researcher
- 2008/01 - 2010/05 University of California, Berkeley Department of Molecular and Cell Biology Ph.D. student
- 2004/08 - 2007/12 University of California, Berkeley Department of Molecular and Cell Biology Graduate student (exchange student)
- 2004/04 - 2004/07 Kyoto University Doctor's Program Graduate student
- 2002/04 - 2004/03 Kyoto University Graduate school of Biostudies, Master's Program Graduate student (Master's program)
- 1998/04 - 2002/03 Kyoto University Department of Human Studies Undergraduate student
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