- 2016 - 2020 Structual study of photo-activated meta state of squid rhodopsin
- 2012 - 2016 Molecular mechanisms in a visual transduction system of invertebrate rhodopsin
- 2012 - 2015 Higher order structure and function of retinal proteins
- 2011 - 2013 無脊椎動物ロドプシンの光活性化機構の解明
- 2010 - 2011 Structural analysis of photo-intermediates of squid rhodopsin
- 2009 - 2011 Structural and functional divergence of rhodopsin super-family
- 2008 - 2009 Gq型G蛋白質共役光受容体のX線構造解析
- 2007 - 2008 X-ray crystallographic study of squid rhodopsin
- 2007 - 2008 Analyses of common structural motif of rhodopsin family
- 2005 - 2006 Four Dimensional Structural Analysis of Biological Ion Pumps
- 2003 - 2004 X-ray Crystallographic Analyses of Retinal Proteins
- 2000 - 2001 生体イオンポンプの反応中間体のX線結晶構造解析
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