- 2021 - 2025 Studies on the pleiotropic functions and transcriptional regulation of gbx homeobox genes in the development of the vertebrate brain.
- 2018 - 2021 Regulatory mechanism of the differentiation of multipotent stem cells in the tailbud and spinal cord development in vertebrate embryos.
- 2015 - 2018 Building new foundation for biological sciences based on transposon-mediated gene trapping
- 2014 - 2017 Studies on the genetic network allowing the integrated regulation of brain regionalization and neurogenesis in vertebrate embryos
- 2012 - 2016 Toward Brain Census: Construction of a method for revealing the memory mechanism at the molecular level
- 2011 - 2013 Mechanism of the development of local organizers in the brain primordium using zebrafish as a model system
- 2008 - 2010 Regulatory mechanism of the formation of the signaling center for regionalization of the brain primordium in vertebrate embryos.
- 2005 - 2007 Molecular mechanism of the formation of the signaling center in the prospective hindbrain of vertebrate early embryos
- 2003 - 2004 Analysis of the gene network patterning the hindhtain in vertebrates.
- 1998 - 2003 脊椎動物中枢神経系における前後軸・背腹軸の成立機構
- 1997 - 1999 Joint Study on Gene Expression in Early Einbryo
- 1997 - 1998 ウニ初期胚における Homeobox 遺伝子の機能の解析
- 1996 - 1996 ウニ上皮成長因子相同ペプチド前駆体遺伝子の初期発生における転写調節機構の解析
- 1991 - 1991 ウニ胚の発生における上皮成長因子ホモローグ(外腸胚形成ペプチド)の機能の解析
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