- 2012/04 - 現在 Doshisha University Graduate School of Brain Science
- 2009/06 - 2012/03 Doshisha University Faculty of Life and Medical Sciences, Department of Medical Life Systems
- 2008/05 - 2009/05 Tokyo Medical and Dental University The Center for Brain Integration Research
- 2008/04 - 2008/04 Tokyo Medical and Dental University Graduate School of Medical and Dental Sciences
- 2008/02 - 2008/03 Tokyo Medical and Dental University Graduate School of Medical and Dental Sciences
- 2004/09 - 2008/02 Tokyo Medical and Dental University Graduate School of Medical and Dental Sciences
- 1997/09 - 2004/08 独マックスプランク生物物理化学研究所 研究員
- 2002/04 - 2004/03 (日本学術振興会海外特別研究員)
- 1994/04 - 1997/03 日本学術振興会特別研究員(DC1)
- 1993/04 - 1997/03 東京大学大学院農学生命科学研究科獣医学専攻博士課程
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