- 2021 - 2024 真核生物のゲノム解析から探るシリカ被殻形態制御の進化と顕生代の古海洋環境
- 2018 - 2022 Origin and evolution of the silica fixation ability of polycystine radiolarians based on molecular phylogenetic analysis.
- 2015 - 2018 Origin of establishment of symbiotic relationship between radiolarians and the symbionts and evolution of the shell components based on the molecular phylogeny and the ultrastructural observations.
- 2011 - 2013 Evolution and origin of the chemical composition of radiolarian test during geologic time scale
- 2007 - 2009 Evolution of the ability to form a siliceous test of protists based on the molecular phylogeny
- 2007 - 2009 A study to develop the program for the environmental learning to foster scientific expression of children. -Through understanding patterns in natural things and phenomena-
- 2003 - 2005 Morphological and molecular evolution of Radiolaria based on the 18S rDNA study
- 2003 - 2005 A study for development of teacher traing program to nurture earth science lireracy through field activities in science.
- 2000 - 2002 Molecular study for symbiosis of the Polycystine radiolarians
- 1998 - 1999 STUDIES OF BIODIVERSITY ON PROTOZOAN ENDOSYMBIOTIC DIATOMS
- 1995 - 1995 西南日本外帯の付加体および付加体上の自亜紀堆積盆形成史の解明-放散虫化石を用いた地質年代・地質構造・堆積(付加)物質量の推定-
- 地質古生物学・付加体地質学
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