- 2020 - 2023 Studies on species biodiversity and evolution of enigmatic parasitic dinoflagellates, Haplozoon spp., which acquired multicellularity.
- 2014 - 2017 Origin of deep-dwelling coccolithophores
- 2012 - 2016 Unexploited habitat of microalgae - investigating biodiversity in the lower photic zone
- 2011 - 2013 Evolutional size change in living and fossil Gephyrocapsa
- 2009 - 2012 Taxonomic investigation on the protozoan fauna of Japan, with special reference to the diversity crisis due to global climate change.
- 2008 - 2012 Studies on life cycles of aquatic parasites, in particular, host-specificities of their infective stages
- 2007 - 2009 Studies on biodiversity and phylogeny of soil protists, with special reference to soil flagellates.
- 2004 - 2006 Studies on diversity and origins of parasitic dinoflagellates.
- 2002 - 2004 Taxonomy and ecology of parasites on zooplankton, in particular, copepods
- 2001 - 2003 Studies on biodiversity and origins of freshwater dinoflagellates.
- 1998 - 1999 PHYLOGENETIC AND TAXONOMIC STUDY OF SUBTERRANEAN WATER PROTISTS.
- 1997 - 1999 Systematic studies on Malaysian marine algae
- 1996 - 1997 Research on the dinoflagellate eyespot-their diversity, division mechanisms and origins-.
- 1994 - 1996 A molecular phylogenetic analysis of the fucalean brown algae based on RFLPs of chloroplast DNA.
- 1994 - 1995 Ultrastructure and ontogeny of eyespots in dinoflagellates : an approach from the perspective of their endosymbiotic origins.
- 1994 - 1995 クロミスタ生物群の細胞特性と系統の研究
- 1991 - 1993 Comparative taxonomy of algae assemblages in mangal environments.
- 1990 - 1990 真核細胞を内部共生体にもつ渦鞭毛藻類の生活環と細胞構造の解析
- 藻類の微細構造、分子系統、分類
- Ultrastructure, molecular phylogeny and taxonomy of algae and protists
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SOCIETY OF EVOLUTIONARY STUDIES, JAPAN
, JAPANESE SOCIETY FOR PLANT SYSTEMATICS
, アメリカ藻類学会(PHYCOLOGICAL SOCIETY OF AMERICA)
, 国際藻類学会(INTERNATIONAL PHYCOLOGICAL SOCIETY)
, THE JAPANESE SOCIETY OF PHYCOLOGY
, 日本植物学会