- 2023 - 2026 Adaptive Strategies and Enabling Conditions of Seasonal Tropical Tree Species Exhibiting Unusual Delayed leaf flushing phenology
- 2017 - 2021 The geohistorical role of mountainous environment causing high diversity of woody plants in Japan
- 2016 - 2020 Evolutionary roles of hybrid zones of the sect. Brachycaryx azaleas in the Fossa Magna Area
- 2013 - 2017 The role of long seed dispersal by large mammals in maintaining biological diversity of forests
- 2012 - 2016 人工林に残された保残帯の面積と配置がハナバチの送粉機能に与える効果の解明
- 2012 - 2015 A study on the hybrid speciation mechanism in plants using genome information
- 2009 - 2011 Studies on effects of the outbreeding depression on reproduction and regeneration of rare tree species grown in wet lands
- 2006 - 2008 Studies on effects of the inbreeding load on reproduction and regeneration of rare tree species grown in wet lands
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