- 2021/04 - National Institute for Environmental Studies Biodiversity Division Senior Researcher
- 2018/04 - 2021/03 National Institute for Environmental Studies Center for Environmental Biology and Ecosystem Studies Senior Researcher
- 2018/05 - 2018/10 Stanford University Department of Biology Research Scholar
- 2013/04 - 2018/03 National Institute for Environmental Studies Center for Environmental Biology and Ecosystem Studies Researcher
- 2010/04 - 2013/03 The Graduate University for Advanced Studies School of Advanced Sciences
- 2007/04 - 2010/03 Kyoto University Graduate School of Agriculture JSPS Postdoctoral Fellow
- 2007/07 - 2008/10 University of Zurich Institute of Plant Biology
- 2006/04 - 2007/03 Research Institute for Humanity and Nature
- 2005/04 - 2006/03 Kyoto University Graduate School of Science JSPS Fellow
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