- 2023 - 2026 Diversity of fern independent gametophytes in Japan and their origin
- 2019 - 2022 Search of new independent gametophytes of ferns in Japan using DNA barcoding
- 2018 - 2021 Analyzing genome constitution of apogamous ferns by much of SNPS data
- 2017 - 2021 Diversity on the flora and traditional use of wild plants in the cultural region of evergreen broadleaf forest
- 2016 - 2019 Genome constitution and reticulate evolution in apogamous ferns
- 2014 - 2017 オシダ属の無配生殖種における網状進化
- 2013 - 2016 Identification of the genetic regions responsible for apogamous reproduction in ferns
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